We know there are microorganisms living some good kms above our heads --- bacteria, fungi, even algae were detected in meteorological samples. What if the unstable mixing of gases that happens in the troposphere and some occasional thunderstorms have cooked up those microbes so they evolved into multicellular organism and then eventually to more complex lifeforms.
I was thinking of some kind of colonial organism like Portuguese man o' war, sea sponges and corals. Tiny polyps connected together to form airborne structures.
Let's call it Atmoid.
Atmoid's biochemistry could adapt to convert atmospheric gases and sunlight into energy and store water in the form of vapor, maybe using solar beams to create a greenhouse effect within its translucent body heating up its inner gases so they become less dense, which means Atmoid is basically a living balloon.
With basically an infinite source of energy, Atmoid can grow to colossal sizes, absorbing most of the sunlight that passes through it and casting a ominous shadow on the equilibrium of the biomes bellow.
Since some tropospheric microbes can work as aerosols and catalyze bio-precipitations (it means bacteria can make it rain), maybe Atmoid can have some weather manipulation abilities.
Catalyzing clouds to create rain and blizzards decreases the temperature in lower regions, which increases the density difference that Atmoid needs to stay afloat, so Atmoid can grow even more without bothering with weight.
Atmoid grows as the sunless world bellow freezes and enters a new ice age.
Atmoid's size eventually circumvents Earth and covers all the sky. No, IT IS THE SKY!
Atmoid now absorbs all the energy that comes from the sun, its single planetary colony now reached level 1 civilization without even known how to make fire!
Claps for Atmoid!
What's the plausibility of such a organism without much handwaving?